180 Dendroculus Locations - Genshin Impact Sumeru Desert 3.1

There are a total of 271 Dendroculi in Genshin Impact, but only 180 Dendroculi are available in Version 3.1 (70 new Dendroculi in the desert). Important note: Make sure you complete the Golden Slumber quest, Dual Evidence, and Afratu’s Dilemma to fully unlock the Sumeru Desert and collect all available Dendroculi.

Location No.01

Teleport to the statue located south of Mawtiyima Forest and head south. Use the Four-Leaf Sigil to teleport and glide east toward the mountain.

Glide toward the Dendroculus.

Location No.02

Return to the same mountain and head southeast. Use the Four-Leaf Sigil to teleport and collect the Dendroculus.

Location No.03

Head northeast and climb the mountain. You can regenerate stamina by climbing toward the yellow flower. Use the Four-Leaf Sigil to teleport and collect the Dendroculus.

Location No.04

Glide northwest and collect the Dendroculus.

Location No.05

Glide northwest from the previous location.

Location No.06

Head south and use the Four-Leaf Sigil to travel faster. The Dendroculus is at ground level.

Location No.07

Teleport to the Mawtiyima Forest west waypoint and glide northeast. The Dendroculus is located in the middle section of the glowing mushroom.

Location No.08

Glide northeast and use the Four-Leaf Sigil. You may have collected this Dendroculus during the Aranyaka quest series.

Location No.09

Teleport to the Mawtiyima Forest southwest waypoint and glide southwest. Enter the cave and explore further to find the Dendroculus on top of a large leaf.

Location No.10

Teleport to the Mawtiyima Forest west waypoint, glide northeast, and descend to ground level. Enter the cave, glide northwest, and use the Four-Leaf Sigil. Perform a Dendro attack on the rock to reveal the Dendroculus.

Location No.11

Teleport to the Mawtiyima Forest west waypoint, glide east, and climb the mountain. Glide west to reach the mid-air Dendroculus.

Location No.12

Teleport to the Palace of Alcazarzaray east waypoint and head northwest. Climb the tree to find the Dendroculus on top of the structure.

Location No.13

Teleport to the Palace of Alcazarzaray west waypoint and head west toward the bouncy mushroom. Look upward and use the Four-Leaf Sigil to collect the Dendroculus.

Location No.14

Glide north from the previous location and collect the Dendroculus floating inside the cave.

Location No.15

Teleport to the Palace of Alcazarzaray west waypoint and head southeast. The Dendroculus is floating beneath the platform.

Location No.16

Teleport to the Palace of Alcazarzaray west waypoint, head west, enter the cave, and explore further. Collect Dendrograna and perform charged attacks on all three floating targets to remove the bubble.

Location No.17

Head southeast to find the Dendroculus on top of the pillar.

Location No.18

Teleport to the Palace of Alcazarzaray south waypoint and head south. The Dendroculus is floating beside the waterfall.

Location No.19

Teleport to Sumeru City west waypoint and head west. The Dendroculus is floating mid-air above a bouncy mushroom.

Location No.20

Teleport to Yazadaha Pool southeast waypoint and head northwest. Hit the Dendro bomb beside the stone to raise it and reveal the Dendroculus.

Location No.21

Head north from the previous location to find the Dendroculus under the bridge.

Location No.22

Teleport to Yazadaha Pool southeast waypoint and head southeast. The Dendroculus is inside a hollow tree trunk.

Location No.23

Teleport to the Gandharva Ville Statue of the Seven and head west.

Location No.24

Teleport to the Gandharva Ville Statue of the Seven and head north. Complete the Aranyaka quest chain if the cave entrance is blocked.

Location No.25

Teleport to the far west waypoint of Vanarana and glide northeast.

Location No.26

Teleport to the Vanarana Statue of the Seven and head north.

Location No.27

Teleport to the Vanarana Statue of the Seven and head west. Climb the mountain and collect the Dendroculus inside the hollow tree trunk.

Location No.28

Teleport to the Vanarana Statue of the Seven and head southwest. Enter the cave filled with Sweet Flowers and collect the Dendroculus inside the house.

Location No.29

Teleport to the Vanarana east waypoint and head northeast. Apply Electro to the bouncy mushroom to reach the Dendroculus.

Location No.30

Head southeast from the previous location and use the Four-Leaf Sigil to swing toward the Dendroculus.

Location No.31

Teleport to the Vanarana east waypoint and head southwest. Continue southwest to find the Dendroculus inside a tent.

Location No.33

Head southwest and use the bouncy mushroom to reach the Dendroculus.

Location No.34

Teleport to the Vanarana southeast waypoint and head northeast.

Location No.35

Head northwest, follow the path, and explore until you find the Dendroculus.

Location No.36

Teleport to the statue west of Yazadaha Pool and glide west. Enter the area above the waterfall and head northeast.

Location No.37

Teleport to the Fane of Ashvattha northwest waypoint and head northwest. Perform a Dendro attack on the rock to reveal the Dendroculus.

Location No.38

Teleport to the Old Vanarana Statue of the Seven and head northeast. Use a Dendro attack to reveal the Dendroculus.

Location No.39

Teleport to the far west waypoint of Vanarana and glide southeast. The Dendroculus is floating at the cave entrance.

Location No.40

Teleport to Sumeru City west waypoint, head west, and climb the tree branch to collect the Dendroculus.

Location No.41

Teleport to the Gandharva Ville Statue of the Seven and head west. Climb the tree branch to find the Dendroculus.

Location No.42

Teleport to Yazadaha Pool southwest waypoint and head north. Apply Electro to the bouncy mushroom to reach the Dendroculus.

Location No.43

Glide east from the previous location and use the bouncy mushroom to collect the Dendroculus.

Location No.44

Head south and collect the mid-air Dendroculus.

Location No.45

Teleport to Pardis Dhyai north waypoint and head west. Use a Dendro attack on the rock to reveal the Dendroculus.

Location No.46

Head north, enter the cave, and explore. Swing to the top of the orange mushroom to collect the Dendroculus.

Location No.47

Teleport to the Fragment of Childhood Dreams domain and reach the higher platform to collect the Dendroculus.

Location No.48

Explore further and use the Four-Leaf Sigils to teleport upward.

Location No.49

Teleport to Old Vanarana southwest waypoint and head east. The Dendroculus is on top of the structure.

Location No.50

Glide northeast and perform a Dendro attack on the rock to reveal the Dendroculus.

Location No.51

Head east to find the Dendroculus on a tree.

Location No.52

Head southeast. The Dendroculus is floating on top of a rock pile.

Location No.53

Head west and remove the bubble by hitting all three targets using Dendrograna.

Location No.54

Teleport to Old Vanarana southwest waypoint and head south. The Dendroculus is on top of a structure.

Location No.55

Teleport to Caravan Ribat northeast waypoint and head west. Perform a Dendro attack on the rock to reveal the Dendroculus.

Location No.56

Teleport to the Fane of Ashvattha south waypoint and head northeast. Apply Electro to the bouncy mushroom to collect the Dendroculus.

Location No.57

Glide west and use the bouncy mushroom to reach the Dendroculus.

Location No.58

Teleport to Caravan Ribat northeast waypoint and head northeast. The Dendroculus is at the cave entrance.

Location No.59

Teleport to Port Ormos northeast waypoint and head northeast. The Dendroculus is on top of a tree.

Location No.60

Glide northeast and interact with all four flowers. Defeat the enemies to remove the bubble.

Location No.61

Head northwest. The Dendroculus is on a tree.

Location No.62

Head northeast from the previous location.

Location No.63

Head east and remove the bubble using Dendrograna.

Location No.64

Head west to find the Dendroculus beside a purple flower.

Location No.65

Head northeast. The Dendroculus is between tree trunks.

Location No.66

Teleport to Devantaka Mountain northeast waypoint and head northeast.

Location No.67

Teleport to Devantaka Mountain northeast waypoint, head northwest, and climb northward.

Location No.68

Teleport to Vimara Village north waypoint and head east. The Dendroculus is beside an Aranara.

Location No.69

Head south and use a Dendro attack on the rock.

Location No.70

Head southeast using the Four-Leaf Sigils. The Dendroculus is inside a broken hut.

Location No.71

Head west to find the Dendroculus at the cave entrance. This is often collected during the Aranyaka quest.

Location No.72

Teleport to Vimara Village southwest waypoint and head west. The Dendroculus is floating mid-air.

Location No.73

Teleport to Vimara Village southwest waypoint and head east. The Dendroculus is floating mid-air.

Location No.74

Teleport to Vimara Village southwest waypoint and head southwest. Apply Electro to the bouncy mushroom.

Location No.75

Head northwest. The Dendroculus is under the bridge.

Location No.76

Continue northeast to find the Dendroculus at a dead end.

Location No.77

Teleport to Devantaka Mountain Statue of the Seven and head south. The Dendroculus is inside a hollow tree trunk.

Location No.78

Continue south and summon a Waverider.

Location No.79

Sail northeast and collect the Dendroculus on the net.

Location No.80

Head northeast from the previous location.

Location No.81

Teleport to Port Ormos northeast waypoint and glide northwest. The Dendroculus floats in front of the giant Ruin Guard.

Location No.82

Glide west. The Dendroculus is inside the Ruin Guard’s hand.

Location No.83

Head northeast to find the Dendroculus on top of a mechanical gear.

Location No.84

Teleport to Devantaka Mountain southeast waypoint and follow the Four-Leaf Sigils southeast. Use the bouncy mushroom.

Location No.85

Teleport to Port Ormos northeast waypoint and head southwest. Play Do–So–La–Mi–Do on the Vintage Lyre to remove the barrier.

Location No.86

Teleport to Port Ormos northwest waypoint and head northeast. The Dendroculus is on top of a cogwheel.

Location No.87

Teleport to Port Ormos northwest waypoint and head northeast. The Dendroculus is on top of a house.

Location No.88

Teleport to Port Ormos northwest waypoint and head southwest. The Dendroculus is inside a hollow tree trunk.

Location No.89

Teleport to Port Ormos northwest waypoint and head west. The Dendroculus is inside a hollow tree trunk.

Location No.90

Teleport to The Dark Valley domain and head southwest. Use a Dendro attack on the rock.

Location No.91

Teleport to Pardis Dhyai Statue of the Seven and head southeast. Use a Dendro attack on the rock.

Location No.92

Head southeast from the previous location.

Location No.93

Head southwest to find the Dendroculus beside the waterfall.

Location No.94

Teleport to Apam Woods southeast waypoint, face north, and climb the tree.

Location No.95

Glide north to collect the Dendroculus at the cave entrance.

Location No.96

Head southwest. Use Electro on the bouncy mushroom, activate the Four-Leaf Sigil, and collect the Dendroculus mid-air.

Location No.97

Teleport to Apam Woods west waypoint and head south. Use a Dendro attack on the rock.

Location No.98

Head northeast and collect the Dendroculus above the waterfall.

Location No.99

Teleport to Apam Woods southeast waypoint and head west. Collect the Dendroculus at the cave entrance.

Location No.100

Teleport to Apam Woods southeast waypoint and head southwest. The Dendroculus is floating above the water.

Location No.101

Teleport to the Ruins of Dahri southeast waypoint and head west. Climb the tree to collect the Dendroculus.

Location No.102

Head northwest and enter the portal to collect the Dendroculus.

Location No.103

Exit the portal, head northeast, and use the bouncy mushroom. Collect the Dendroculus from the tree branch.

Location No.104

Drop down and head east using the Four-Leaf Sigils.

Location No.105

Teleport to the Ruins of Dahri southeast waypoint and head northeast. Use a Dendro attack on the rock.

Location No.106

Head southeast and collect the Dendroculus beneath the wooden platform.

Location No.107

Continue southeast and use a Dendro attack on the rock.

Location No.108

Teleport to Apam Woods west waypoint, head southeast, enter the cave, and use the bouncy mushroom.

Location No.111

Teleport to Aaru Village waypoint. Climb the building to collect the Dendroculus on the rooftop.

Location No.112

Head northwest, swing upward, and glide to collect the Dendroculus.

Location No.113

Plunge down and head south.

Location No.114

Swing upward and head southwest. The Dendroculus is floating beneath the platform.

Location No.115

Teleport to the Abdju Pit domain and head west.

Location No.116

Teleport to Khaj-Nisut west waypoint, head north, and climb the structure.

Location No.117

Glide east, enter the ruin, go right, and plunge down. The Dendroculus is on top of the chest.

Location No.118

Return along the path to find a Dendroculus hidden behind a broken wall.

Location No.119

Teleport to Khaj-Nisut west waypoint, head east, and enter the ruin. Turn left, descend three levels, turn right, and collect the Dendroculus.

Location No.120

Teleport to Khaj-Nisut southeast waypoint and glide northwest toward the top of the structure.

Location No.121

Glide southwest, enter the ruin, descend through the hole, and find the Dendroculus behind the statue.

Location No.122

Teleport to Khaj-Nisut southeast waypoint and glide northeast. The Dendroculus is on top of a rock.

Location No.123

Teleport to Dar al-Shifa northwest waypoint and head southeast. The Dendroculus is inside a ground hole.

Location No.124

Remove the breakable rocks while progressing through the underground path to reach the Dendroculus.

Location No.125

Teleport to Dar al-Shifa southwest waypoint and glide west toward the tree.

Location No.126

Teleport to Abdju Pit east waypoint and head southeast. The Dendroculus is on the elevator.

Location No.127

Use the platform and elevator mechanics to reach the Dendroculus.

Location No.128

Find another Dendroculus northwest after reaching the higher platform.

Location No.129

Navigate staircases and elevators to reach the Dendroculus on the higher level.

Location No.130

Follow the elevator and platform sequence to reach the Dendroculus atop the statue.

Location No.131

Explore the large room, use the elevator, and collect the Dendroculus behind the statue.

Location No.132

Teleport to Sobek Oasis northeast waypoint and climb the structure.

Location No.133

Teleport to Valley of Dahri northeast waypoint and swing upward to collect the Dendroculus.

Location No.134

Teleport to the Dune of Elusion east waypoint and head northeast.

Location No.135

Glide east toward the top of the Fatui camp.

Location No.136

Teleport to Valley of Dahri Statue of the Seven, head north, and enter the hole.

Location No.137

Teleport to Valley of Dahri southwest waypoint and head northeast. Swing upward to reach the Dendroculus.

Location No.138

Enter the ruin, explore to the end, and climb the tree branch to collect the Dendroculus.

Location No.139

Teleport to Valley of Dahri southwest waypoint and head southeast. Swing upward to the tree branch.

Location No.140

Continue southeast to find the Dendroculus on the giant ruin.

Location No.141

Teleport to Valley of Dahri northwest waypoint and head south. Complete Afratu’s Dilemma if blocked.

Location No.142

Climb the root and collect the Dendroculus behind the cogwheel.

Location No.143

Teleport to Valley of Dahri Statue of the Seven and head south. Enter the cave to find the Dendroculus.

Location No.144

Teleport to Valley of Dahri Statue of the Seven and head east. Enter the cave and collect the floating Dendroculus.

Location No.145

Teleport to the Dune of Elusion east waypoint and head west. The Dendroculus is on top of a tree.

Location No.146

Teleport to the Dune of Elusion waypoint and head northwest. Reach the top of the pyramid.

Location No.147

Head south and light all three Pyro monuments to collect the Dendroculus.

Location No.148

Teleport to the Dune of Magma waypoint and head southwest. Follow the ruin puzzle sequence to collect the Dendroculus.

Location No.149

Exit the ruin, activate the fan, and glide upward to collect the Dendroculus.

Location No.150

Place the blue pyramid, complete the sandglass challenge, and collect the Dendroculus below.

Location No.151

Head northeast to collect the Dendroculus on top of the pillar.

Location No.152

Teleport to the Mausoleum of King Deshret northwest waypoint and glide north.

Location No.153

Complete the ruin puzzle sequence to unlock the elevator and collect the Dendroculus.

Location No.154

Teleport to the Dune of the Carouses northwest domain and swing upward to collect the Dendroculus.

Location No.155

Teleport to the Dune of Carouses northeast waypoint. The Dendroculus is behind three wooden crates.

Location No.156

Teleport to the Dune of Carouses northeast waypoint, enter the ruin, and collect the Dendroculus.

Location No.157

Teleport to Khemenu Temple Statue of the Seven and head northwest. The Dendroculus is floating above the platform.

Location No.158

Glide west along the wind current to collect the Dendroculus.

Location No.159

Teleport to Khemenu Temple Statue of the Seven and head south. The Dendroculus is on top of a statue.

Location No.160

Head southwest from the previous location.

Location No.161

Continue west to find the Dendroculus beside the staircase.

Location No.162

Teleport to the Mausoleum of King Deshret east waypoint and head west. Turn right three times.

Location No.163

Return to the main room, enter the southern door, and turn left twice.

Location No.164

Teleport to the Dune of Magma waypoint, head west, and drop down. The Dendroculus is floating mid-air.

Location No.165

Teleport to the Dune of Magma north waypoint and head northeast.

Location No.166

Teleport to the Dune of Magma north waypoint and explore west. Swing to the top of the glowing mushroom.

Location No.167

Teleport to the Mausoleum of King Deshret southwest waypoint and head northwest.

Location No.168

Descend to the lower level and head north to find the Dendroculus on a statue.

Location No.169

Head southwest and explore until you see a tree with the Dendroculus.

Location No.170

Teleport to the Mausoleum of King Deshret north waypoint and head south. The Dendroculus is on a pillar.

Location No.171

Continue south and climb the invisible platform beneath the Dendroculus.

Location No.172

Teleport to the Mausoleum of King Deshret north waypoint and head south. Find the Dendroculus behind the hologram pyramid.

Location No.173

Teleport to the Mausoleum of King Deshret south waypoint, enter the blue pyramid, and descend carefully.

Location No.174

Teleport to the Mausoleum of King Deshret southeast waypoint and navigate the ruin to find the Dendroculus.

Location No.175

Teleport to Khemenu Temple west waypoint, head west, and enter the door on the right.

Location No.176

Teleport to Khemenu Temple west waypoint and explore south. The Dendroculus is on a boat.

Location No.177

Navigate downward through the rooms to find the floating Dendroculus near the broken bridge.

Location No.178

Explore west until reaching wooden structures. Climb and glide to the Dendroculus.

Location No.179

Teleport to Khemenu Temple west waypoint and navigate the ruin puzzle to collect the Dendroculus.

Location No.180

Teleport to the Dune of Elusion waypoint and complete the ruin puzzle to reach the final Dendroculus.

If you have collected all Dendroculi from Version 3.0 and 3.1, you should reach Statue of the Seven Level 7 with 23 extra Dendroculi remaining.